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Q.

Gymnosperm leaves have adapted to withstand extremes in temperature, humidity, and  [[1]] 

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answer is WIND.

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Detailed Solution

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Concept: The leaves of gymnosperm are highly adapted to tolerate temperature, humidity, and wind extremes. 
The leaves of conifers are needle like which limit the surface area. 
The thick cuticle and sunken stomata also aid in water conservation.
Hence,  the wind is the correct answer.

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