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Q.

Shuttle vectors:

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a

Can replicate in both eurkaryotic cell and E. coli 

b

Contain two types of origin of replication and selectable marker genes, one for eukaryotic cell and the other for E. coli

c

Includes YEp (Yeast Episomal Plasmid) and modified Ti-plasmid

d

Is related to all of the above options 

answer is D.

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Detailed Solution

Yeast episomal plasmids and modified Ti plasmids are shuttle vectors which can multiply in two different hosts, have two ori sites and selectable markers one for eukaryotes and another one for prokaryotes. 

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Shuttle vectors: