The principle of competitive exclusion was stated by

The principle of competitive exclusion was formulated by Russian ecologist Georgy Gause in the 1930s, based on his experiments with two species of Paramecium. It states that two species that compete for the same resources cannot coexist indefinitely.
(1) Charles Darwin is known for his theory of evolution by natural selection.
(3) MacArthur is known for his work on niche theory and island biogeography. He observed resource partitioning in
Warbler species.
(4) Verhulst and Pearl are known for their work on population growth models.