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Through his long-term ecosystem experiments using outdoor plots, David Tilman showed that

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loss of key species is a more serious threat to the ecosystem.

when resources are limited, the competitively superior species will eliminate the other species.

increased diversity contributed to higher productivity.

species richness increased with increasing explored area, but only up to a limit.

answer is A.

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Tilman found that plots with more species showed less year-to-year variation in total biomass. He also showed that in his experiments, increased diversity contributed to higher productivity.

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