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a
originate in the lumen of vessels
b
characterise the sapwood
c
are extensions of xylem parenchyma cells into vessels
d
are linked to the ascent of sap through xylem vessels.
answer is C.
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Detailed Solution
Tyloses are balloon shaped extensions of xylem parenchyma into vessels. They arise during secondary growth. When pressure built up on older secondary xylem (heartwood) due to successive rings of sapwood, xylem parenchyma cells intrude in vessels and block it. Every structure developed in lumen of vessels is not tyloses.