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Jan Ingenhousz - hypothesized that plants restore to the air whatever breathing animals and burning candles remove.

Priestley - showed that it is only the green part of the plants that could release oxygen.

Julius von Sachs - demonstrated that photosynthesis is essentially a light-dependent reaction.

T.W Engelmann - described the first action spectrum of photosynthesis.

answer is 4.

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Using a prism T.W Engelmann split light into its spectral components and then illuminated a green alga, Cladophora, placed in a suspension of aerobic bacteria. The bacteria were used to detect the sites of O2 evolution. He observed that the bacteria accumulated mainly in the region of blue and red light of the split spectrum. A first action spectrum of photosynthesis was thus described. It resembles roughly the absorption spectra of chlorophyll a and b.

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