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Bergmann's Rule

By Swati Singh

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Updated on 14 Apr 2025, 18:21 IST

Have you ever realised why polar bears are large-sized while their tropical relatives, like sun bears, are much smaller? Or why people living in colder regions are heavily built than those from the warmer areas? Let us explain these interesting observations with the help of Bergmann's Rule?

This rule will focus on how animals adapt to different climates and environments. In this article, we will discuss what Bergmann's Rule is, why it takes place, a few examples, as well as some exceptions to this rule, and finally we will rule out common misconceptions surrounding the rule.

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What is Bergmann's Rule?

It is an eco-geographical principle that correlates the temperature of an environment and the ratio of body surface to weight in warm-blooded animals. It explains that a larger body size of an organism living in colder regions of the world helps in preserving heat by minimizing heat loss while those living in warmer regions don’t need to do that and hence are comparatively smaller in size.

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This principle was first proposed by a German biologist, Carl Bergmann, in 1847. However, this principle is only applied to endothermic animals such as mammals and birds.

Why Does Bergmann’s Rule Occur?

The principle is based on thermoregulation - the ability of an organism to maintain its body temperature. Larger animals have a smaller surface-area compared to their body volume, which helps them retain heat more efficiently in cold regions. Whereas smaller animals, that have a larger surface-area-to-volume ratio, lose heat faster. This makes them better suited for warmer climates.

Bergmann's Rule

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To understand Bergmann’s Rule better, let's break it down into key points:

  • Surface Area Volume: Larger animals have a proportionally smaller surface area relative to their volume, reducing heat loss in cold climates.
  • Energy Conservation: A larger body mass allows animals to store more energy and endure harsh conditions with fewer food resources.
  • Heat Dissipation: In contrast, smaller animals in warm climates have a higher surface area relative to their volume, helping them release excess body heat.
  • Evolutionary Adaptation: Over generations, natural selection favours traits that enhance survival in specific environmental conditions.

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Examples & Applications of Bergmann’s Rule

Bergmann’s Rule is widely observed across various animal species. Let’s explore some real-world examples:

1. Bears (Ursidae Family)

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  • Polar bears (Arctic regions) are the largest bear species, adapted to extreme
  • Brown bears (temperate regions) are slightly smaller but still
  • Sun bears (tropical forests) are the smallest, suited for warm

2. Deer and Moose

  • Moose in northern latitudes (Canada, Scandinavia) are much larger than their deer relatives in warmer regions.

3. Human Populations

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  • Inuit and other Arctic populations generally have stockier builds compared to people from tropical regions, which aligns with Bergmann’s Rule.

Exceptions to Bergmann’s Rule

Although Bergmann’s Rule is common, not all animals follow it. Some exceptions include:

  • Marine mammals: Whales and dolphins don’t strictly follow this rule because their body size is influenced more by swimming efficiency than temperature.
  • Some birds: Tropical birds like toucans have large bodies, even in warm
  • Small mammals: Some desert rodents, like kangaroo rats, are relatively large compared to others, but this helps them conserve water rather than regulate temperature.

These exceptions show that other factors, like environment and diet, also influence body size.

Bergmann’s Rule is often confused with Allen’s Rule, which states that animals in colder regions tend to have shorter appendages (ears, tails, limbs) to minimize heat loss. While both rules deal with thermoregulation, Bergmann’s Rule focuses on overall body size, whereas Allen’s Rule addresses limb and extremity length.

Conclusion

Bergmann’s Rule provides valuable insights into how animals adapt to different climates by regulating body size for thermal efficiency. From Arctic polar bears to tropical sun bears, this principle explains fascinating variations in nature.

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Understanding Bergmann’s Rule not only enriches our knowledge of evolution and ecology but also has practical implications for conservation, wildlife management, and even predicting biological responses to climate change.

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Bergmann's Rule FAQs

Does Bergmann’s Rule apply to all animals?

No, Bergmann’s Rule mainly applies to warm-blooded animals (endotherms) like mammals and birds. Cold-blooded animals (ectotherms) do not follow this rule as their body temperature depends on external conditions.

Are there exceptions to Bergmann’s Rule?

Yes, some species do not follow Bergmann’s Rule due to factors like diet, predation, and migration patterns. Additionally, human activity and climate change are influencing natural size variations.

How does Bergmann’s Rule relate to climate change?

As global temperatures rise, some species may experience changes in body size. Studies suggest that warming climates are leading to smaller body sizes in some birds and mammals.

Can Bergmann’s Rule be applied to extinct species?

Yes, palaeontologists use Bergmann’s Rule to study size variations in prehistoric animals, such as Ice Age mammals, which were generally larger than their modern relatives.

Is Bergmann’s Rule a strict law of biology?

No, it is a general trend rather than a strict law. While it explains many observations, exceptions exist due to complex ecological interactions.